Bowser returns as a villain for this sequel. It is a planetoid in the shape of Mario's head which uses a map screen, with a word scheme similar to the ones in previous Mario games: Super Mario Bros. The new method of traveling through space is the Starship Mario, built by Lubba. However, the Comet Observatory has been eliminated as Mario's means of accessing the galaxies. Star Bits, as well as suits such as the Bee Mushroom and other mechanics of Super Mario Galaxy reappeared in the game. It is like Super Mario World and also has some Super Mario World themes. Miyamoto stated that most of what didn't make it into Super Mario Galaxy would appear in Super Mario Galaxy 2. There is also a switch that slows/stops time, a rotating item block similar to a die that Mario can use his spin attack on-gaining whatever is rolled on the block, winged Question Blocks and more. Also found are two new suits called Rock Mario and Cloud Mario the former turns Mario into a rock ball that rolls around hitting enemies, and the latter lets Mario make clouds to climb. New items include a drill head, used to perforate planets. Yoshi can gain new powers by grabbing special items, such as the Dash Pepper, which lets him run up steep slopes, the Blimp Fruit, which causes him to float, and the Bulb Berry, which allows him to find hidden platforms.

The most notable feature in Super Mario Galaxy 2 is Yoshi, whom Mario can ride on.

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